Physiology and Behaviour of Animal Suffering (UFAW Animal Welfare)

Suffering is a state of mind that is difficult to measure and analyse in human beings and considerably more so in animals. It is related to the environment in which we live and our physical and mental states. Understanding the physiology of suffering in animals is crucial in assessing animal welfare. Written by an expert in applied welfare aspects of physiology, this book is the first to address the physiological aspects of suffering in animals. It explores the different causes of suffering – physical discomfort, thirst and hunger, the responses in the body that lead to suffering and it offers insight into how suffering can be managed. The second book in a major new animal welfare series Draws together information that is scattered across the literature Written for the specialist and non-specialist alike Includes colour pictures This book is part of the UFAW/Wiley-Blackwell Animal Welfare Book Series. This major series of books produced in collaboration between UFAW (The Universities Federation for Animal Welfare), and Wiley-Blackwell provides an authoritative source of information on worldwide developments, current thinking and best practice in the field of animal welfare science and technology. For details of all of the titles in the series see www.wiley.com/go/ufaw. Physiology and Behaviour of Animal Suffering......Page 5 Contents......Page 7 Foreword......Page 11 Preface......Page 13 1.3 When Can We Stop Worrying about Animal Suffering?......Page 15 1.4 Recognising Suffering in Animals......Page 21 1.5 Can Animals Go Mad?......Page 22 1.6 What Constitutes Animal Suffering?......Page 23 1.7 Conclusions......Page 24 2.1 Stress Physiology......Page 26 2.2 Stress-related Disorders......Page 31 2.3 Restraint Stress......Page 33 2.5 Stress-induced Seizures and Fits......Page 35 3.1 Introduction......Page 36 3.2 Anxiety......Page 37 3.3 Experimental Models......Page 39 3.4 Fear......Page 40 3.6 Phobias......Page 42 3.7 Panic......Page 43 3.8 The Role of the Amygdala......Page 44 3.9 Losing One’s Mother......Page 45 4.1 Anhedonia......Page 46 4.2 Depression......Page 47 4.3 Social Isolation in the Newborn......Page 48 4.4 Sensory Deprivation in Early Life......Page 49 4.6 Social Isolation and Barren Environments......Page 50 4.7 Stereotypies and Neurotic Behaviours......Page 51 4.9 Post-traumatic Stress Disorder......Page 54 4.10 Sleep Disorders......Page 56 4.11 Weaning......Page 58 4.12 Physiology of Emotions......Page 59 5.1 Aggression......Page 60 5.2 Overcrowding and Confinement......Page 65 5.3 Discomfort......Page 68 6.1 Overexertion......Page 70 6.2 Endurance Riding......Page 71 6.3 Horse-racing Injuries......Page 72 6.4 Greyhound Racing Injuries......Page 75 6.5 Migration......Page 77 7.1 Cold Discomfort and Pain......Page 78 7.2 Skin Freezing and Chilblains......Page 79 7.3 Hypothermia......Page 80 7.5 Hypothermia and Cold Survival......Page 81 7.6 Cold in Combination with Starvation......Page 83 7.7 Cold-induced Analgesia......Page 84 8.1 Heat Stress......Page 86 8.2 Heat Intolerance......Page 87 8.3 Some Species Differences......Page 88 8.4 Pain......Page 89 8.5 Burns and Scalds......Page 90 9.1 Thirst and Dehydration......Page 97 9.3 Osmotic Stress......Page 100 9.4 Hunger......Page 101 9.5 Underfeeding......Page 102 9.6 Emaciation......Page 103 9.7 Inappropriate Diets......Page 105 9.9 Force-feeding and Overeating......Page 106 10.1 The Value of Pain......Page 108 10.2 Pain Associated with Trauma......Page 109 10.3 Ways in Which Animals Express Pain......Page 113 10.4 Pain Pathways and Consciousness......Page 116 10.5 Cortical Regions......Page 120 10.6 Applied Neurology of Pain......Page 121 10.7 Pain in a Given Context......Page 133 11.1 Injuries in Selected Body Regions and Tissues......Page 145 11.2 Some Common Causes of Injury......Page 147 11.3 Intended or Avoidable Causes of Trauma......Page 156 11.4 Types of Injury......Page 186 12.1 Is Suffering an Inevitable Consequence of Being Ill?......Page 197 12.2 Do the Behaviours Expressed During Sickness Serve a Purpose?......Page 199 12.3 Cytokines and Sickness Behaviours......Page 200 12.4 Cancer......Page 201 12.5 Stress and Immune Function......Page 202 12.8 Hazards of Improving Disease Control......Page 205 12.9 Diseases Used for Controlling Pests......Page 206 13.2 Vomiting and Retching......Page 207 13.3 Gut Pain......Page 209 13.4 Diarrhoea......Page 211 13.6 Stress and the Gut......Page 212 13.7 Gastrointestinal Ulcers......Page 213 14.1 Wartime Poisons......Page 215 14.2 Environmental Toxicants......Page 216 14.3 Vertebrate Pesticides......Page 217 15 Respiratory System......Page 221 15.1 Asphyxia......Page 222 15.2 Breathlessness......Page 223 15.3 Carbon Dioxide Inhalation......Page 225 15.4 Drowning......Page 226 15.5 Pulmonary Oedema......Page 228 15.6 Hypoxia......Page 229 15.7 Altitude Sickness......Page 231 15.8 Decompression......Page 232 15.10 Asthma and Allergies......Page 233 15.12 Signs of Respiratory Distress......Page 234 15.13 Agonal Gasping......Page 236 16 Dying......Page 237 16.1 Euthanasia......Page 238 16.2 Decapitation......Page 240 16.4 Death from Brain Injury......Page 241 16.5 Recognising Insensibility and Brain Death......Page 243 References......Page 246 Abbreviations......Page 272 Index......Page 274 Colour plate sections falls after page 148......Page 162

Suffering is a state of mind that is difficult to measure and analyse in human beings and considerably more so in animals. It is related to the environment in which we live and our physical and mental states. Understanding the physiology of suffering in animals is crucial in assessing animal welfare.

Written by an expert in applied welfare aspects of physiology, this book is the first to address the physiological aspects of suffering in animals. It explores the different causes of suffering – physical discomfort, thirst and hunger, the responses in the body that lead to suffering and it offers insight into how suffering can be managed.

• The second book in a major new animal welfare series
• Draws together information that is scattered across the literature
• Written for the specialist and non-specialist alike
• Includes colour pictures

This book is part of the UFAW/Wiley-Blackwell Animal Welfare Book Series. This major series of books produced in collaboration between UFAW (The Universities Federation for Animal Welfare), and Wiley-Blackwell provides an authoritative source of information on worldwide developments, current thinking and best practice in the field of animal welfare science and technology. For details of all of the titles in the series see www.wiley.com/go/ufaw.

1. Introduction; 2. Stress; 3. Anxiety and fear; 4. Emotional numbness and deprivation; 5. Aggression, overcrowding and discomfort; 6. Exercise; 7. Cold; 8. Heat and burns; 9. Thirst and hunger; 10. Pain; 11. Trauma; 12. Sickness and disease; 13. Digestive system; 14. Poisoning; 15. Respiratory system; 16. Dying. "Written by an expert in applied welfare aspects of physiology, this book is the first to address the physiological aspects of suffering in animals. It explores the different causes of suffering - physical discomfort, thirst and hunger - the responses in the body that lead to suffering and it offers insight into how suffering can be managed."--BOOK JACKET